diff --git a/includes/bootstrap.inc b/includes/bootstrap.inc
index 8c043378e284..48b287546a6b 100644
--- a/includes/bootstrap.inc
+++ b/includes/bootstrap.inc
@@ -832,7 +832,9 @@ function drupal_page_is_cacheable($allow_caching = NULL) {
  *   The name of the bootstrap hook we wish to invoke.
  */
 function bootstrap_invoke_all($hook) {
-  foreach (module_list(TRUE, TRUE) as $module) {
+  // _drupal_bootstrap_page_cache() already loaded the bootstrap modules, so we
+  // don't need to tell module_list() to reset its bootstrap list.
+  foreach (module_list(FALSE, TRUE) as $module) {
     drupal_load('module', $module);
     module_invoke($module, $hook);
   }